This volume describes a 3.6 million-years-old partial skeleton of Australopithecus afarensis from the Woranso-Mille, central Afar, Ethiopia. This specimen is the first adult partial skeleton to be recovered since Lucy’s (A.L. 288-1) discovery in 1974. It is older than Lucy by 400,000 years and sheds light on the paleobiology of earlyAustralopithecus afarensis, particularly the morphology of the shoulder girdle and thoracic shape, which are thus far poorly understood and actively debated. The fauna associated with the partial skeleton tells us enormously about Au. afarensis paleoecology and give us another piece of the puzzle regarding habitat availability and use for Au. afarensis outside the Hadar region where it has been well-known for the last four decades.