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This volume is number four in a series of proceedings volumes from the International Symposia on Fractals in Biology and Medicine in Ascona, Switzerland which have been inspired by the work of Benoît Mandelbrot seeking to extend the concepts towards the life sciences. It highlights the potential that fractal geometry offers for elucidating and explaining the complex make-up of cells, tissues and biological organisms either in normal or in pathological conditions, including the structural changes that occur in tumours. It helps develop the concepts, questions and methods required in research on fractal biology and natural phenomena and to evidence the pitfalls of a too simplistic application of these principles in investigating topical subjects of biology and medicine. It discusses present and future applications of fractal geometry, bringing together cellular and molecular biology, engineering, mathematics, physics, medicine and other disciplines and allowing an interdisciplinary vision. The book should be of interest to researchers and students from molecular and cell biology, biomedicine, biomathematics, analytical morphology, immunology and neurology who are interested in the combination of mathematics and life sciences.