Genesis Debt (The Genesis Project‪)‬

In the spectacular sequel to Genesis 2, the defeat of the alien matriarch, Link-soo-shan, has united the highly-evolved species known as the Gnathe with the human race. But peace is to be short-lived…
Mankind began a new existence on Jupiter millions of years after the sun expanded to become a red giant and eradicated all life on Earth. But they were not alone on their new home. The Gnathe, creatures with an incredibly aptitude for genetic engineering, had already established their civilization there and some of them did not welcome their new neighbours.
Led by Link-soo-shan, the humans narrowly escaped being made extinct for good. With the help of winged beasts known as Terra-raptors, the humans deposed Link-soo-shan and formed a new alliance with the Gnathe.
In Genesis Debt the science-fiction adventure continues as new technologies are discovered and the new colonies attempt to uncover where the Gnathe came from and whether they can travel backwards to save Earth from its catastrophic fate.
But in the shadows the rumour of Link-soo-shan's return grows and it is clear that she has her own plans for revenge on the humans and their allies…